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ID-FRONTIERS for March 16-22, 2003

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Messages are displayed in the order they were received.
 Subject From Date  Time 
 Franklin's Gull molt questions  Martin Reid   Sun, 16 Mar 2003  5:50am 
 Gull help requested (again)  Nick Komar   Sun, 16 Mar 2003  6:40am 
 Iceland/Thayer's Gull BNA  Jean Iron   Sun, 16 Mar 2003  10:20am 
 Re: Iceland/Thayer's Gull BNA  Phil Pickering   Sun, 16 Mar 2003  11:11am 
 Re: Franklin's Gull molt questions  Tony Leukering   Sun, 16 Mar 2003  11:53am 
 Re: Iceland/Thayer's Gull BNA  Alvaro Jaramillo   Sun, 16 Mar 2003  1:20pm 
 Re: Iceland/Thayer's Gull BNA  Millington/BIS   Sun, 16 Mar 2003  2:04pm 
 Spizella sparrow near Elma(photos) (fwd)  Ian Paulsen   Sun, 16 Mar 2003  5:04pm 
 Re: Iceland/Thayer's Gull BNA  Alvaro Jaramillo   Sun, 16 Mar 2003  6:10pm 
 Re: Iceland/Thayer's Gull BNA  Anthony L. Lang  Sun, 16 Mar 2003  7:37pm 
 "Kumlien's" x Thayer's 'intergrades' and IDing glaucoides ICGUs  Michael Retter   Sun, 16 Mar 2003  9:42pm 
 Laughing x Ring-billed hybrid  Brian Boldt   Sun, 16 Mar 2003  10:18pm 
 Re: Iceland/Thayer's Gull BNA  Alain Foss=?ISO-8859  Sun, 16 Mar 2003  10:34pm 
 Re: Laughing x Ring-billed hybrid  Alan Wormington   Mon, 17 Mar 2003  11:04am 
 Varied Thrush Puzzle  Angus Wilson   Mon, 17 Mar 2003  10:32pm 
 Ageing & Sexing Varied thrushes  Julian Hough   Tue, 18 Mar 2003  9:53am 
 Re: Varied Thrush Puzzle  Angus Wilson   Tue, 18 Mar 2003  9:55am 
 Re: Varied Thrush Puzzle  Julian Hough   Tue, 18 Mar 2003  11:15am 
 Re: Varied Thrush Puzzle  Tony Leukering   Tue, 18 Mar 2003  11:17am 
 Re: Varied Thrush Puzzle  Les Chibana   Tue, 18 Mar 2003  2:00pm 
 Re: Varied Thrush Puzzle  Mike Patterson   Tue, 18 Mar 2003  3:38pm 
 Re: Varied Thrush Puzzle  Angus Wilson   Tue, 18 Mar 2003  7:46pm 
 VA Beach Gull  John Puschock   Tue, 18 Mar 2003  9:28pm 
 Identification of Brant - Variability and hybridization  Angus Wilson   Tue, 18 Mar 2003  11:37pm 
 Re: Varied Thrush Puzzle  Nathan Hentze   Wed, 19 Mar 2003  12:31am 
 Re: VA Beach Gull  Nick Rossiter   Wed, 19 Mar 2003  7:51am 
 Plumage variation in Varied Thrush  Julian Hough   Wed, 19 Mar 2003  10:27am 
 Re: Plumage variation in Varied Thrush  Matt Kenne   Wed, 19 Mar 2003  11:35am 
 Re: Winter Laughing Gulls  Ian Mclaren   Wed, 19 Mar 2003  12:45pm 
 Re: Varied Thrush Puzzle  Tony Leukering   Wed, 19 Mar 2003  4:15pm 
 Re: Winter Laughing Gulls  Tony Leukering   Wed, 19 Mar 2003  4:22pm 
 Re: Winter Laughing Gulls  Matt Kenne   Wed, 19 Mar 2003  7:28pm 
 Re: VA Beach Gull  Martin Reid   Thu, 20 Mar 2003  5:05am 
 RE : VA Beach Gull  =?iso-8859-1?Q?Pierr  Thu, 20 Mar 2003  6:52am 
 Re: Winter Laughing Gulls  D. Heindel  Thu, 20 Mar 2003  8:24am 
 Re: Winter Laughing Gull  Ian Mclaren   Thu, 20 Mar 2003  8:41am 
 Re: Winter Laughing Gull  Alvaro Jaramillo   Thu, 20 Mar 2003  9:42am 
 Winter LAGU  Gavin Bieber   Thu, 20 Mar 2003  10:01am 
 Re: VA Beach Gull  Peter Adriaens   Thu, 20 Mar 2003  10:50am 
 VA Beach Gull  Nick Rossiter   Thu, 20 Mar 2003  1:24pm 
 Re: Winter Laughing Gull  Floyd Hayes   Thu, 20 Mar 2003  2:10pm 
 Fwd: Re: Winter Laughing Gull  Alvaro Jaramillo   Fri, 21 Mar 2003  12:07am 
 RE : VA Beach Gull  =?iso-8859-1?Q?Pierr  Fri, 21 Mar 2003  2:47am 
To use email addresses replace '(AT)' with '@'.
This is done to confuse the spam 'bots.


[ << | >> | ^^ ] Subject: Franklin's Gull molt questions From: Martin Reid <upupa(AT)AIRMAIL.NET> Date: 16 Mar 2003 5:50am Dear all, Can anyone point me to published works that discuss in detail the molt of Franklin's Gull? My limited library contains few details beyond the idea (from Grant's GULLS) that FRGUs undergoes a complete pre-migration molt in the Spring and Fall (except for the partial first prebasic molt.) This is repeated in Sibley's Field Guide, but I do not have any other references that go into more detail. I ask because I recently noticed that some of the FRGUs arriving just now in Fort Worth, Texas seem to have two generations of primaries; see these examples: http://www.martinreid.com/frgu03.html - is this an aberrant suspension of molt, or a normal partial molt? (and if the latter, is it a hallmark of a particular age?) Also FYI here's an adult FRGU that completely lacked white eye crescents: http://www.martinreid.com/frgu02.html Good Birding, Martin Reid Fort Worth, Texas mailto:upupa(AT)airmail.net http://www.martinreid.com
[ << | >> | ^^ ] Subject: Gull help requested (again) From: Nick Komar <quetzal65(AT)ATTBI.COM> Date: 16 Mar 2003 6:40am My apologies to the list for supplying a hyperlink that was non-functional. This one should work: http://www.birdinghawaii.co.uk/WELCOMEPAGE.htm. Click on the link for News, Events and Announcements, and then click on the gull photo to see the series of photos of this interesting gull in Oahu. Nick Komar Fort Collins CO ----- Original Message ----- From: "Nick Komar" <quetzal65(AT)attbi.com> To: <BIRDWG01(AT)listserv.arizona.edu> Cc: "Peter Donaldson" <PDonaldson(AT)hawaii.rr.com>; "birding hawaii" <birdinghawaii(AT)yahoo.com> Sent: Saturday, March 08, 2003 5:44 PM Subject: Gull help requested > Photos of a first-basic gull that I saw in February in Oahu (Hawaii) are > posted at > http://photos.groups.yahoo.com/group/hawaiibirding/vwp?.dir=/2003+Photograph > s&.src=gr&.dnm=Komar+Gull+1.jpg&.view=t&.done=http%3a//photos.groups.yahoo.c > om/group/hawaiibirding/lst%3f%26.dir=/2003%2bPhotographs%26.src=gr%26.view=t > (clicking on this link brings up the first of three photos). > > I suspect "Vega" Gull (Larus argenteus vegae) because of the bright white > rump in constrast with a dark mantle. I have no experience with this > species. Can a Vega Gull expert comment on whether other features support > the identification? Vega Gull is extrememly rare in Oahu (possibly never > reported). > > I'd be interested in opinions as to the identity of this gull. Thanks. > > Nick Komar > Fort Collins CO > > > > > > >
[ << | >> | ^^ ] Subject: Iceland/Thayer's Gull BNA From: Jean Iron <jeaniron(AT)SYMPATICO.CA> Date: 16 Mar 2003 10:20am Account number 699 of the Iceland Gull (nominate glaucoides and kumlieni) and Thayer's Gull (monotypic) in The Birds of North America by Richard Snell arrived last week. There is much new data from Snell's original field work and his examination of adult Iceland and Thayer's Gull specimens collected from throughout the breeding range. Here are some highlights in "quotes" and my comments. 1. Introduction: Snell says on page 2 "My studies, heretofore unpublished, are suggestive of continuous phenotypic variation from darkest to lightest extremes in plumage of adults across the breeding range. Where breeding ranges of kumlieni and thayeri overlap (e. Baffin I., Southampton I., Digges Sound), there is no evidence of assortative mating: gulls as dark or darker than the type of thayeri bred with others much lighter than the type of kumlieni, including birds lacking visible wing-tip melanism. Based on this, I believe only 1 species should be recognized with all taxa placed under Iceland Gull." 2. Thayer's Gull: On page 3, "least melanistic thayeri, including some specimens from Eureka, Ellesmere I., are lighter than dark kumlieni." There is no information where these pale-winged thayeri from Ellesmere spend the winter. "Iris color highly variable, including in western part of breeding range of thayeri..." It is now generally agreed that eye color is extremely variable from dark to clear yellow in both thayeri and kumlieni, but averages darker in thayeri. On page 9, the character on which thayeri is most distinct is mantle melanism, though there is broad overlap with kumlieni." 3. Kumlien's Gull: On page 8, "Melanism of irides variable, extent of speckling in irides variable; and no evidence of significant correlation between melanism of irides and patterns on primary-feathers of adults on e. Baffin I...Wing-tips of breeding adults range from patternless to relatively restricted patterns shaded gray to dark gray with partial white apical spots (Table 1). On average, mantles and P10 webs darker gray than nominate glaucoides, but there is overlap between subspecies. Plumage and mensural characters of kumlieni overlap both nominate glaucoides and thayeri." Table 1 on page 9 referred to above shows the supporting data from 7 geographic regions across the breeding range of the three taxa from Greenland to Banks and Finlayson Islands in Canada. Figure 6 on page 6 is a map showing the 7 regions used in this account and shows the locations of the type specimens of kumlieni and thayeri. The type specimen of glaucoides was from Iceland, but is not extant. 4. Nominate Iceland Gull: On page 8, "Although most sw. and e. Greenlandic adults (i.e., well south of Thule region) lack wing-tip melanism, many nominate glaucoides possess gray wing-tips and light patterning on primary-feathers...Two e. Greenlandic chicks captured Jul 1964, and reared in captivity, had markings on inner and outer webs of P10 when fully adult...if viewed in the south as winter migrants, these birds would doubtless be considered kumlieni. Near Nuuk, sw. Greenland, birds with patterned wing-tips were relatively numerous in 2001: one was documented on a nest 16 Jun...On average, male and female glaucoides have the smallest tarsi and culmens, but there is overlap in mensural characters with kumlieni." On page 12, "Most adults wintering in Iceland have traces of melanism on one or more primaries." The above suggests introgression with kumlieni. 5. Figure 3 on page 10: This illustration by David Beadle shows the range of variation in 18 wing-tip patterns of thayeri, kumlieni and glaucoides collected in the breeding range of Canada and Greenland. The illustration includes the type specimens of thayeri and kumlieni. I believe that the wing-tip drawing of the type specimen of Thayer's is not what most people would call a typical Thayer's in the field, especially on the west coast. The first drawing shows a dark thayeri wing-tip with a mirror on only P10 that is essentially the same as many NA Herring Gulls. Note that Snell does not suggest that Thayer's should be merged with Herring Gull. This dark extreme shows that wing-tip pattern and melanism varies in different directions in thayeri. Snell states on page 9, "The type specimens for kumlieni and thayeri may represent points in the clinal continua of glaucoides, kumlieni, and thayeri rather than exemplars of discrete taxa..." You can buy a copy of Snell's account of Iceland and Thayer's in The Birds of North America, number 699, from Buteo Books http://www.buteobooks.com/bna.html I look forward to the ongoing discussions of these fascinating gulls. Ron Pittaway Minden and Toronto, Ontario E-mail: jeaniron(AT)sympatico.ca Jean Iron President Ontario Field Ornithologists 9 Lichen Place Toronto ON M3A 1X3 416-445-9297 e-mail: jeaniron(AT)sympatico.ca www.ofo.ca
[ << | >> | ^^ ] Subject: Re: Iceland/Thayer's Gull BNA From: Phil Pickering <philliplc(AT)HARBORSIDE.COM> Date: 16 Mar 2003 11:11am Recognizing kumlieni as an intergrade is one thing, but (asking in ignorance of any genetic studies that have been done) does clinal variation and interbreeding automatically make nominate glaucoides and thayeri one species? How does this situation differ from Western and Glaucous-winged, other than that introgression is (perhaps) geographicaly more progressed? Cheers, Phil > 1. Introduction: Snell says on page 2 "My studies, heretofore unpublished, > are suggestive of continuous phenotypic variation from darkest to lightest > extremes in plumage of adults across the breeding range. Where breeding > ranges of kumlieni and thayeri overlap (e. Baffin I., Southampton I., > Digges Sound), there is no evidence of assortative mating: gulls as dark or > darker than the type of thayeri bred with others much lighter than the type > of kumlieni, including birds lacking visible wing-tip melanism. Based on > this, I believe only 1 species should be recognized with all taxa placed > under Iceland Gull."
[ << | >> | ^^ ] Subject: Re: Franklin's Gull molt questions From: Tony Leukering <GreatGrayOwl(AT)AOL.COM> Date: 16 Mar 2003 11:53am ----INCLUDING text/plain MIME SECTION---- Hi all: John Vanderpoel and I, while John was preparing his Small Gulls ID video, noted that most of the Franklin's Gulls (FRGU) in Alternate I that he had video of showed a similar pattern of primary retention to that of many buteos, specifically that they retained a few of the outermost juvenal primaries. John and I discussed it at length, but, unfortunately, he did not have enough examples to make any sweeping statements so, I believe, left that factoid out of the video. Now, we did not note any such pattern in adults, so.... Sincerely, Tony Leukering Brighton, CO ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Iceland/Thayer's Gull BNA From: Alvaro Jaramillo <chucao(AT)ATTBI.COM> Date: 16 Mar 2003 1:20pm At 10:17 AM 3/16/2003 -0800, Phil Pickering wrote: >Recognizing kumlieni as an intergrade is one thing, but (asking in >ignorance of any genetic studies that have been done) does clinal >variation and interbreeding automatically make nominate glaucoides >and thayeri one species? How does this situation differ from >Western and Glaucous-winged, other than that introgression >is (perhaps) geographicaly more progressed? > >Cheers, > >Phil Phil, This doesn't answer your question, but interbreeding (hybridization) and clinal variation are two different situations. In most cases a cline occurs over a substantial geographic area, and change over that area is gradual. Most geographic variation in Red-winged Blackbirds and Savannah Sparrows (to give examples of species that have been studied extensively) is clinal. These clines make it impossible for one to separate most populations as discrete entities from each other. So most named subspecies of both of these species are not workable as they are specific points on a cline rather than distinct populations that differ diagonsably from the next population over. The typical cline occurs within a species. When you have a hybrid zone (such as in Western/Glaucous-winged gulls) the variation can be termed a "step cline" since you have two levels (the parent species) and a narrow area where the two are intermediate (the step). But these are not clines in the way that one traditionally thinks of a cline: Classical cline - from state A to state B: --------------------------- B ----------------------------- --------------------------- A ----------- Step cline (hybrid zone) from state A to state B ------------------------------------------------------- B -- --- --- A ----------------------------- The x-axis is geographical distance, y axis some morphological feature, or composite of morphology. In a very simplistic way to look at complex problems, if two "types" or "taxa" exhibit a clear cline in morphology such as the top example, then it suggests that the two are not separate. They are not on evolutionary distinct trajectories, they in fact are part of the same thing. More than likely these two should be treated as the same species. If differentiation between populations A and B are not possible due to the completeness of the cline, then you can't even give them separate subspecies status. Now if genetic work shows that A and B are not each other's closest relatives and they are behaving in this manner, that is a problem. The answers on what to do with it are more muddy. It is clear now that species don't always just arrive and remain, but they may be 'swallowed' up by hybridizing with another species, or perhaps a new species could even arise from a hybrid population, or at least a population that has genes it incorporated from historical hybridization. Biologists refer to this generally as 'reticulation', and the true pattern of divergence in many species may be much more like a reticulated web, than the nice little branching trees we always picture. In the hybrid zone, or step cline example (the bottom one) the two taxa may very well be different species. They are retaining their overal morphological (and genetic) differences, but only in a limited part of their range are they mixed up in this hybrid situation. If the two forms are not each other's closest relatives (sister species) then it makes sense to retain them as separate species, otherwise your phylogeny of the group becomes muddled with species that do not properly reflect the history of the group. Available evidence, mainly genetic, has shown that Western Gull and Glaucous-winged Gull are not sister species, the same goes for Bullock's and Baltimore Orioles. Its tougher to decided what to do when the two taxa are sisters, such as in Myrtle and Audubon's Warbler which show a step cline in morphology in the limited area where they hybridize. Some sister taxa in this situation are considered separate species such as Townsend's Warblers and Hermit Warblers, but not in others such as the Yellow-rumped Warbler example. Some of this difference is just due to inconsistencies in treatment by taxonomic bodies as understanding of speciation and new techniques that allow one to look at gene flow and genetic differentiation have evolved. Some of it is also due to what is published and known, versus yet unknown. Factors that need to be considered are what are the levels of gene flow between the populations, are the clines stable?, does the genetic data suggest that the two taxa have been separate for a long period of time (genetic distances between populations), etc. But to get back to the Thayer's x Iceland situation, I think that part of what is happening is that it is unclear which one of these two patterns the group falls into. Richard Snell has data that is strongly suggestive that it is the clinal pattern. Clines in the Arctic Islands are really difficult to deal with however. If you look at the geography of the place there is a large north-south gradient as well as east-west gradient where things can happen, and the pattern of variation in this complex seems to differ depending on where you are in the Arctic. Thayer's are not just west of 'kumlien's', they are also north of them. If you look at a globe and look at the Arctic from above, there are probably breeding areas of Thayer's that are closer geographically to nominate Iceland than they are to Kumlien's (would need to pull the globe myself and figure out if this is the case, but it is certainly not unlikely). Apart from the messy geography, these are also islands and islands function in their own peculiar manner. So who knows, there may be classic clines within taxa here, as well as hybrid zones. I think two things have to happen before any of this can be resolved. First of all, the data set on geographic variation in morphology of this group needs to be published and assessed. Also we need to clarify if Thayer's and Iceland are sister taxa or not, and I recall that the latest gull work by Trochet suggest they are not. Steve Howell and Bruce Mctavish's recent paper in Alula is interesting in that it shows that most adult Kumlien's are separable from most Thayer's based on field studies of large numbers of each on the wintering grounds. In other words, there is little overlap in plumage pattern between the two, although the differences are subtle. I think they make a good argument that this is the case. What is also true, looking at their data, is that Kumlien's Gull is rather variable. When you think about this, its odd. Sure there is a lot of variability in gulls, particularly when they are young, but as adults the variability shown by most taxa is not nearly as much as that shown by Kumlien's Gull. Why is it so variable? There has to be a reason. The suggestion that Kumlien's is of hybrid origin between Thayer's and Iceland fits neatly as an answer to this. Perhaps its not all that dissimilar from the Western and Glaucous-winged Gull situation in this respect. Imagine that "Olympic Gull" or "Puget Sound Gull" had been described as a different entity, and we believed it was a different entity. It would make our understanding of the relationships between Western and Glaucous-winged Gull a lot less clear. cheers Alvaro Alvaro Jaramillo Biologist San Francisco Bay Bird Observatory P.O. Box 247 Alviso, CA 95002 (408)-946-6548 http://www.sfbbo.org/ chucao(AT)attbi.com
[ << | >> | ^^ ] Subject: Re: Iceland/Thayer's Gull BNA From: Millington/BIS <sales(AT)BIRDINGWORLD.CO.UK> Date: 16 Mar 2003 2:04pm Hi Alvaro's comments are most lucid and very helpful. The recent Alula article goes one or two steps further than Alvaro mentions, and suggests that Kumlien's Gull, although variable, is c95% diagnosable and may possibly merit species status. Furthermore, there is the suggestion that 'pale-winged' Thayer's may result from thayeri and glaucoides occasionally hybridising to the north of kumlieni (hypothetically) Certainly, there seems a very solid case for thayeri and glaucoides being diagnosable units (species), with kumlieni (possibly consisting of two forms, say the authors....) being either a hybrid swarm or a recognisable taxon. The evidence for this is substantial, as a high proportion of each form that can be seen in the wintering areas where, as with some other gulls (& brent geese and many ducks etc), the appearance and variation of the majority of birds is far easier to assess than on the breeding grounds. If the situation is clinal as Snell states, where do all the (thousands of?) confusing 'intergrades' winter? Currently they remain just hypothetical entities cheers Richard sales(AT)birdingworld.co.uk (Birding World, Books for Birders & Birdline) Bird Information Service, Stonerunner, Coast Road, Cley next the Sea, Holt, Norfolk, NR25 7RZ, UK Tel. 44 (0) 1263 741139 Fax. 741173 Website www.birdingworld.co.uk -----Original Message----- From: Alvaro Jaramillo <chucao(AT)ATTBI.COM> To: <BIRDWG01(AT)listserv.arizona.edu> Date: 16 March 2003 20:25 Subject: Re: [BIRDWG01] Iceland/Thayer's Gull BNA >At 10:17 AM 3/16/2003 -0800, Phil Pickering wrote: >>Recognizing kumlieni as an intergrade is one thing, but (asking in >>ignorance of any genetic studies that have been done) does clinal >>variation and interbreeding automatically make nominate glaucoides >>and thayeri one species? How does this situation differ from >>Western and Glaucous-winged, other than that introgression >>is (perhaps) geographicaly more progressed? >> >>Cheers, >> >>Phil > >Phil, > > This doesn't answer your question, but interbreeding (hybridization) >and clinal variation are two different situations. In most cases a cline >occurs over a substantial geographic area, and change over that area is >gradual. Most geographic variation in Red-winged Blackbirds and Savannah >Sparrows (to give examples of species that have been studied extensively) >is clinal. These clines make it impossible for one to separate most >populations as discrete entities from each other. So most named subspecies >of both of these species are not workable as they are specific points on a >cline rather than distinct populations that differ diagonsably from the >next population over. The typical cline occurs within a species. When you >have a hybrid zone (such as in Western/Glaucous-winged gulls) the variation >can be termed a "step cline" since you have two levels (the parent species) >and a narrow area where the two are intermediate (the step). But these are >not clines in the way that one traditionally thinks of a cline: > >Classical cline - from state A to state B: > > --------- ------------------ >B > ----------------------------- > --------------------------- >A ----------- > >Step cline (hybrid zone) from state A to state B > ------------------------------------- ------------------ >B > -- > --- > --- >A ----------------------------- > >The x-axis is geographical distance, y axis some morphological feature, or >composite of morphology. > >In a very simplistic way to look at complex problems, if two "types" or >"taxa" exhibit a clear cline in morphology such as the top example, then it >suggests that the two are not separate. They are not on evolutionary >distinct trajectories, they in fact are part of the same thing. More than >likely these two should be treated as the same species. If differentiation >between populations A and B are not possible due to the completeness of the >cline, then you can't even give them separate subspecies status. Now if >genetic work shows that A and B are not each other's closest relatives and >they are behaving in this manner, that is a problem. The answers on what to >do with it are more muddy. It is clear now that species don't always just >arrive and remain, but they may be 'swallowed' up by hybridizing with >another species, or perhaps a new species could even arise from a hybrid >population, or at least a population that has genes it incorporated from >historical hybridization. Biologists refer to this generally as >'reticulation', and the true pattern of divergence in many species may be >much more like a reticulated web, than the nice little branching trees we >always picture. > >In the hybrid zone, or step cline example (the bottom one) the two taxa may >very well be different species. They are retaining their overal >morphological (and genetic) differences, but only in a limited part of >their range are they mixed up in this hybrid situation. If the two forms >are not each other's closest relatives (sister species) then it makes sense >to retain them as separate species, otherwise your phylogeny of the group >becomes muddled with species that do not properly reflect the history of >the group. Available evidence, mainly genetic, has shown that Western Gull >and Glaucous-winged Gull are not sister species, the same goes for >Bullock's and Baltimore Orioles. Its tougher to decided what to do when the >two taxa are sisters, such as in Myrtle and Audubon's Warbler which show a >step cline in morphology in the limited area where they hybridize. Some >sister taxa in this situation are considered separate species such as >Townsend's Warblers and Hermit Warblers, but not in others such as the >Yellow-rumped Warbler example. Some of this difference is just due to >inconsistencies in treatment by taxonomic bodies as understanding of >speciation and new techniques that allow one to look at gene flow and >genetic differentiation have evolved. Some of it is also due to what is >published and known, versus yet unknown. Factors that need to be considered >are what are the levels of gene flow between the populations, are the >clines stable?, does the genetic data suggest that the two taxa have been >separate for a long period of time (genetic distances between populations), >etc. > >But to get back to the Thayer's x Iceland situation, I think that part of >what is happening is that it is unclear which one of these two patterns the >group falls into. Richard Snell has data that is strongly suggestive that >it is the clinal pattern. Clines in the Arctic Islands are really difficult >to deal with however. If you look at the geography of the place there is a >large north-south gradient as well as east-west gradient where things can >happen, and the pattern of variation in this complex seems to differ >depending on where you are in the Arctic. Thayer's are not just west of >'kumlien's', they are also north of them. If you look at a globe and look >at the Arctic from above, there are probably breeding areas of Thayer's >that are closer geographically to nominate Iceland than they are to >Kumlien's (would need to pull the globe myself and figure out if this is >the case, but it is certainly not unlikely). Apart from the messy >geography, these are also islands and islands function in their own >peculiar manner. So who knows, there may be classic clines within taxa >here, as well as hybrid zones. I think two things have to happen before any >of this can be resolved. First of all, the data set on geographic variation >in morphology of this group needs to be published and assessed. Also we >need to clarify if Thayer's and Iceland are sister taxa or not, and I >recall that the latest gull work by Trochet suggest they are not. Steve >Howell and Bruce Mctavish's recent paper in Alula is interesting in that it >shows that most adult Kumlien's are separable from most Thayer's based on >field studies of large numbers of each on the wintering grounds. In other >words, there is little overlap in plumage pattern between the two, although >the differences are subtle. I think they make a good argument that this is >the case. What is also true, looking at their data, is that Kumlien's Gull >is rather variable. When you think about this, its odd. Sure there is a lot >of variability in gulls, particularly when they are young, but as adults >the variability shown by most taxa is not nearly as much as that shown by >Kumlien's Gull. Why is it so variable? There has to be a reason. The >suggestion that Kumlien's is of hybrid origin between Thayer's and Iceland >fits neatly as an answer to this. Perhaps its not all that dissimilar from >the Western and Glaucous-winged Gull situation in this respect. Imagine >that "Olympic Gull" or "Puget Sound Gull" had been described as a different >entity, and we believed it was a different entity. It would make our >understanding of the relationships between Western and Glaucous-winged Gull >a lot less clear. > >cheers > >Alvaro > >Alvaro Jaramillo >Biologist >San Francisco Bay Bird Observatory >P.O. Box 247 >Alviso, CA 95002 >(408)-946-6548 >http://www.sfbbo.org/ >chucao(AT)attbi.com
[ << | >> | ^^ ] Subject: Spizella sparrow near Elma(photos) (fwd) From: Ian Paulsen <ipaulsen(AT)KRL.ORG> Date: 16 Mar 2003 5:04pm HI: Any comments? Ian Paulsen Bainbridge Island, WA, USA ipaulsen(AT)krl.org A.K.A.: "Birdbooker" "Rallidae all the way" ---------- Forwarded message ---------- Date: Sun, 16 Mar 2003 14:45:21 -0800 From: Ruth Sullivan <GODWIT(AT)worldnet.att.net> To: tweeters(AT)u.washington.edu Subject: Spizella sparrow near Elma(photos) Hello Tweets, My mother and I have placed 4 photos of the unidentified Spizella sparrow that we observed yesterday near Elma,hoping we may get some feedback on the possible identity of the bird between either an abbarent Clay-colored Sparrow or a hybrid between a Chipping and Clay-colored Sparrow. We have placed the 4 photos at our yahoo site at: http://photos.yahoo.com/godwit2003/. The photos labeled sparrow1-3 were taken by mother at very close range and are of the best quality and the photo labeled sparrow4 was taken by me(Patrick) and is of less quality,but perhaps we may still get some feedback from anyone that may give us an opinion in open or privately. Nevertheless,it is a very intersting sparrow,even if it turns out to be a hybrid. Thank you for any assistance, Ruth and Patrick Sullivan godwit(AT)worldnet.att.net
[ << | >> | ^^ ] Subject: Re: Iceland/Thayer's Gull BNA From: Alvaro Jaramillo <chucao(AT)ATTBI.COM> Date: 16 Mar 2003 6:10pm At 09:01 PM 3/16/2003 +0000, Millington/BIS wrote: >Hi > >The recent Alula article goes one or two steps further than Alvaro mentions, >and suggests that Kumlien's Gull, although variable, is c95% diagnosable and >may possibly merit species status. > >Certainly, there seems a very solid case for thayeri and glaucoides being >diagnosable units (species), with kumlieni (possibly consisting of two >forms, say the authors....) being either a hybrid swarm or a recognisable >taxon. Thanks to Richard for giving us a more detailed summary of the Alula article, which is worth a read. In fact all three of Steve Howell's (and collaborators) Alula papers (or was it more?) in the last few years are definitely worth a read. I think that while according to the latest paper there is a low degree of overlap between thayeri and kumlieni based on the features they looked at, this does not necessarily mean there is not a large area of interbreeding, and as Richard's last sentence notes the possibility that this is a hybrid swarm is still there. So while the paper clarifies how to diagnose (ID) the two entities, it gets us no closer to answering the question of how many species are involved here. It also depends on what flavour of species you are most attached to. The high degree of variability within Kumlien's still needs to be explained. >The evidence for this is substantial, as a high proportion of each form that >can be seen in the wintering areas where, as with some other gulls (& brent >geese and many ducks etc), the appearance and variation of the majority of >birds is far easier to assess than on the breeding grounds. Yes and no, but I agree with the overall point that Richard (and the Alula paper makes). The information gathered from wintering birds is extremely useful. It would have been next to impossible to sample this many birds in the breeding range so quickly and so cheaply. However the same information from the breeding grounds would be much more meaningful and could get to the crux of the problem of how many species are involved, by looking at wintering populations you can't do this without making a great number of assumptions, which could be wrong. Only one wintering area for kumlien's was looked at, is it representative of all of the east coast? How would it compare to the Great Lakes? >If the situation is clinal as Snell states, where do all the (thousands of?) >confusing 'intergrades' winter? Currently they remain just hypothetical >entities I bet Great Lakes and St. Lawrence River birds are a bit different, but there sample sizes will be much lower. My guess is that some birds that are considered intergrades by Snell would be considered within the range of variation in kumlien's by Howell and McTavish. How you interpret these intergrade plumages depends on the context, its different when you can see what else is present at the breeding colony for example. Also how you define diagnosibility depends on what you were measuring, and how you analysed these measurements. Howell and McTavish set up their data to be looked at graphically, but there is no statistical test shown that gives you data on the confidence one can have on the diagnosability of these taxa. Basically, how much slop does the data have? What are the confidence limits of the numbers? We don't know. The graphical representation lacks a sample size for Iceland Gulls, was one or a 1000 birds measured? This has bearing on how you interpret the data. They plot proportions against estimated Kodak Values of wingtip darkness, but these are averaged estimates (scores of 7-10, become an 8.5 on the plot). This averaging can lead to a cleaner looking data set than what was really there, and it makes me wonder why they didn't just plot the wingtip scores rather than convert them to kodak values. Finally, and I get back to this variability within Kumlien's, the pattern of the data is important. You have two end points (Iceland and Thayer's) which clearly differ from each other, and show low (non-existent in Iceland here, but no sample size given) amounts of variation within each taxon. Then the intermediate taxon is much more variable than either end point taxon, and at least some individuals overlap with each end point taxon. This is exactly the pattern that you would expect in a hybrid swarm, and exactly the pattern that you would not expect if these were three nice little species going along their own business. Please don't interpret this as a dismissal of what Howell and McTavish have found. Their data is extremely useful, its well worth reading (necessary for any gull aficionado and gull researcher) and they make some solid arguments here. Its great work, and like any good bit of work it asks more questions than it answers. regards Alvaro Jaramillo Biologist San Francisco Bay Bird Observatory P.O. Box 247 Alviso, CA 95002 (408)-946-6548 http://www.sfbbo.org/ chucao(AT)attbi.com
[ << | >> | ^^ ] Subject: Re: Iceland/Thayer's Gull BNA From: "Anthony L. Lang" <tlang(AT)ROGERS.COM> Date: 16 Mar 2003 7:37pm ----- Original Message ----- From: "Millington/BIS" <sales(AT)BIRDINGWORLD.CO.UK> To: <BIRDWG01(AT)listserv.arizona.edu> Sent: Sunday, March 16, 2003 4:01 PM Subject: Re: [BIRDWG01] Iceland/Thayer's Gull BNA > If the situation is clinal as Snell states, where do all the (thousands of?) > confusing 'intergrades' winter? Currently they remain just hypothetical > entities In my experience here around Lake Ontario, the wingtip patterns of putative _kumlieni_ adults (as viewed in flight) are very variable. Perhaps we have the whole range of variation from _thayeri_ through _kumlieni_ to _glaucoides_ right here. There are certainly a few well-documented _thayeri_ known from our area. Now if those Thayer's/Iceland birds would hold still long enough we could have a good look at them and document this variation. However, this variation doesn't tell us much unless we know where each one hatched. Regarding whether classical clinal variation makes _thayer_ and _kumlieni_ one species, in such a case we couldn't separate the birds into diagnosable groups so we would be forced to treat them all as one species (both phylogenetic species and biological species). However, we wouldn't be able to tell whether the cline arose because of hybridization after secondarily coming into contact. ======================================================= Anthony L. Lang #22-6050 Bidwell Trail Mississauga, Ontario, Canada L5V 1V6 tel: 905-812-5513 tlang(AT)rogers.com "Marshall McLuhan got it wrong. It isn't the medium that is the message. It's the marketing." -- Rick Mercer =======================================================
[ << | >> | ^^ ] Subject: "Kumlien's" x Thayer's 'intergrades' and IDing glaucoides ICGUs From: Michael Retter <mretter(AT)HOTMAIL.COM> Date: 16 Mar 2003 9:42pm "If the situation is clinal as Snell states, where do all the (thousands of?) confusing 'intergrades' winter? Currently they remain just hypothetical entities." I can't account for thousands, but I may be able to do so for dozens or even hundreds. Ever since I have been looking at winter gulls here in Illinois, I have been seeing birds I call Thayer's X "Kumlien's" intergrades. First let's define Thayer's X "Kumlien's" intergrade. If Kumlien's is a hybrid swarm between thayeri and glaucoides, then these so-called intergrades are merely dark-end (more thayeri-like) hybrids ("backcrosses"). In this case the term doesn't really have much meaning. BUT, if kumlieni is a good taxon (about which I have serious doubts), then 'Thayer's X "Kumlien's" intergrade' would seem to be a good name for these birds which seem too light for thayeri but too dark for kumlieni. I see individuals that I call intergrades regularly in Illinois. For example, here are my totals this winter from a few days of gulling, at Starved Rock Lock and Dam, LaSalle Co, IL . . . THAYER'S GULL 36+ (7+ 1w, 2 2w, 2 3w, 25+ ad.) 16 Feb. 12+ (3 1w, 1 3w, 1 4w, 7+ ad) 10 Feb. 4 (1w, 1 2w, 2 3w, 15+ ad) 11 Feb. 9+ (2 1w, 1 3w, 6+ ad) 8 Feb. 12+ (2 1w, 1 2w, 1 3w, 8+ ad) 28 Jan. 2 (2 ad) 25 Jan. KUMLIEN'S GULL 3+ (3+ ad) 16 Feb. 3 (1 1w, 2 ad) 10 Feb. 5 (1 1w, 4 ad) 11 Feb. 4+ (1 1w, 1 2w, 2+ ad) 8 Feb. 2 (2 ad) 28 Jan. 3 (3 ad) 25 Jan. INTERGRADES 3 (3 ad) Feb. MR. 5 (1 1w, 4 ad) 10 Feb. 6 (1 3w, 5 ad) 11 Feb. 2 (2 ad) 8 Feb. 2 (2 ad) 25 Jan. This winter I saw just as many intergrades as I did "good" Kumlien's Gulls. "Common knowledge" among Illinois birders seems to be that these birds are out there and are not rare. I distinctly remember the first time I saw such an individual. The bird, an adult, flew past, and I asked "was that a Thayer's or an Iceland?" One of the most experienced observers in the state at the time replied, "Good question. Those are the ones we try to forget we saw." He went on to explain that he sees a few each season. Others there agreed. Unfortunately I can't say that others across the country and the globe would not call what I term intergrades either light thayeri or dark kumlieni, or both. I was able to get pictures of one such bird on 25 Jan 2003, and 3 photos are available for viewing at http://photos.yahoo.com/bc/mlretter/lst?.dir=/other+Starved+Rock+gulls&.src=ph&.view=t Marked "possible intergrade", all 3 photos are of the same bird. Unfortunately, they are all shots of the bird at rest. When the wings were spread they showed an almost perfect thayeri wing pattern, but the primary tip markings were not black. They were a very dark slate gray. The subterminal dark bars on P-8 and P-9 were thinner than I expect on thayeri, but perhaps not out of the range of variation for that taxon. The mantle was slightly lighter than those of the adjacent Herring Gulls. Its eye was amber, not brown, and not yellow, for what that's worth. Other individuals I have called intergrades are similar to this bird; some are not. Some are like Kumlien's with (usually) dark mantles. They have the restricted markings in the primaries like Kumlien's, but the markings are jet black. Someday I hope to know the true status and identification of these birds, but until then, 'intergrade' is the best I can do. Perhaps that desigination is right; perhaps it is not. Any comments would be greatly appreciated. Also at the same URL mentioned above, are 3 pics of a Iceland Gull that looked like a glaucoides. From what we could tell, the wingtips were pure white. The eye however, was amber, not yellow. Is this enough to eliminate glaucoides? Again, any commentary is appreciated. ---------- Michael L. P. Retter Bloomington, McLean Co, IL mlretter(AT)yahoo.com 309.824.7317 see some of my drawings at: http://www.iwu.edu/~mretter/drawings.htm >From: "Anthony L. Lang" <tlang(AT)ROGERS.COM> >Reply-To: "Anthony L. Lang" <tlang(AT)ROGERS.COM> >To: BIRDWG01(AT)listserv.arizona.edu >Subject: Re: [BIRDWG01] Iceland/Thayer's Gull BNA >Date: Sun, 16 Mar 2003 21:37:36 -0500 > >----- Original Message ----- >From: "Millington/BIS" <sales(AT)BIRDINGWORLD.CO.UK> >To: <BIRDWG01(AT)listserv.arizona.edu> >Sent: Sunday, March 16, 2003 4:01 PM >Subject: Re: [BIRDWG01] Iceland/Thayer's Gull BNA > > > > If the situation is clinal as Snell states, where do all the (thousands >of?) > > confusing 'intergrades' winter? Currently they remain just hypothetical > > entities > >In my experience here around Lake Ontario, the wingtip patterns of putative >_kumlieni_ adults (as viewed in flight) are very variable. Perhaps we have >the whole range of variation from _thayeri_ through _kumlieni_ to >_glaucoides_ right here. There are certainly a few well-documented >_thayeri_ known from our area. Now if those Thayer's/Iceland birds would >hold still long enough we could have a good look at them and document this >variation. However, this variation doesn't tell us much unless we know >where each one hatched. > >Regarding whether classical clinal variation makes _thayer_ and _kumlieni_ >one species, in such a case we couldn't separate the birds into diagnosable >groups so we would be forced to treat them all as one species (both >phylogenetic species and biological species). However, we wouldn't be able >to tell whether the cline arose because of hybridization after secondarily >coming into contact. > >======================================================= >Anthony L. Lang >#22-6050 Bidwell Trail >Mississauga, Ontario, Canada >L5V 1V6 >tel: 905-812-5513 >tlang(AT)rogers.com > >"Marshall McLuhan got it wrong. It isn't the medium that is the message. >It's the marketing." -- Rick Mercer >======================================================= _________________________________________________________________ STOP MORE SPAM with the new MSN 8 and get 2 months FREE* http://join.msn.com/?page=features/junkmail
[ << | >> | ^^ ] Subject: Laughing x Ring-billed hybrid From: Brian Boldt <bboldt(AT)EXCELCOMM.COM> Date: 16 Mar 2003 10:18pm This is a multi-part message in MIME format. ----INCLUDING text/plain MIME SECTION---- I'm a new member to the list but have been following on Jack Siler's = site for quite a while; I was prompted to finally join by the wonderful = ongoing Thayer's/Kumlein's thread. I found this bird today and thought I would post to the group as it = makes a nice comparison to Martin Reid's Franklin's hybrid of a while = ago: http://birds.excelcomm.com/gullpage/hybrid.asp Brian Boldt / Milwaukee, WI ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Iceland/Thayer's Gull BNA From: Alain Foss=?ISO-8859-1?B?6Q==?= Angers 49 F <alfosse(AT)WANADOO.FR> Date: 16 Mar 2003 10:34pm Le 17·3·03 2:09, Alvaro Jaramillo à chucao(AT)ATTBI.COM a écrit : > So while the paper clarifies how to > diagnose (ID) the two entities, it gets us no closer to answering the > question of how many species are involved here. In YÉSOU P., 2002. Trends in systematics. Systematics of Larus argentatus-cachinnans-fuscus complex revisited. Dutch Birding, 24 (5) : 271-298. the author thinks kumlieni is a hybrid between the 2 species thayeri and glaucoides. Best regards. •============================================• Alain Fossé,LPO Anjou,Montreuil-Juigné,France 47° 31' 34" N, 0° 35' 43" W <alfosse(AT)wanadoo.fr> My digiscoping <http:/www.digimages.info/> My French List of the Birds of the World <http://membres.lycos.fr/listoiseauxmonde/> Web Site LPO Anjou<http://www.lpo-anjou.org/> •============================================•
[ << | >> | ^^ ] Subject: Re: Laughing x Ring-billed hybrid From: Alan Wormington <wormington(AT)JUNO.COM> Date: 17 Mar 2003 11:04am Several years ago (around March 10th), I observed an adult Laughing X Ring-billed Gull here at Point Pelee (Ontario). Does anyone know if a compilation has been done on this hybrid type re occurrence dates and locations? I can provide the exact details of my sighting, if someone is interested. Alan Wormington Leamington, Ontario
[ << | >> | ^^ ] Subject: Varied Thrush Puzzle From: Angus Wilson <wilsoa02(AT)MED.NYU.EDU> Date: 17 Mar 2003 10:32pm Last night I posted two pictures of a Varied Thrush that was discovered by Al Wollin at Hempstead Lake State Park on Long Island, New York on Saturday afternoon and glimpsed by a small crowd of local birders the following morning. Several correspondents have expressed surprise at my identification of this bird as an adult female. The surprise comes from the fact that the bird was heard singing (brief snatches) on several occasions. http://www.oceanwanderers.com/NYVariedThrush.html The identification as a female is based primarily on the absence of a strong breast band. According to the few sources I can muster (Pyle 1997, Clement 2000 and George 2002), males always show a well-defined but not necessarily complete, breast band. Looking carefully at the photos, which I admit are a little over-exposed, there may be a hint of darker color across the breast. However, this is far weaker than the literature would indicate for males (hatching year or older) and for that matter, weaker than on many adult females. Also supportive of a female are the pale orange tones of the under parts, whitish belly and lower breast and the presence of some whitish patches across the breast. I should add that most of the popular field guides were pretty disappointing on this topic, most failing to explain, let alone illustrate, how first year birds compare to adults. In terms of aging, I feel much less confident. I wrote adult female on the web page (grasping to explain the singing) but this might easily be incorrect. Julian Hough has been thinking about this topic more carefully, in response to a female that is present in Connecticut, and perhaps he can chime in for himself. The relative degree of brown suffusing the dark gray areas on the crown, mantle and upper-side of the tail are key, supplemented with detail of molt and wear. Unfortunately, the very distant and 'from below' views of the Long Island bird make these attributes difficult to assess. So the big question is why is the bird singing? I haven't found any mention in the literature to suggest that female Varied Thrushes sing on a regular basis. If it does occur, one might expect it to be most common in older, more mature, females (elevated testosterone levels?). Luke George mentions that both males and females utter two types of call during aggressive encounters with other birds. These are a 'vree' (with or without a trill) or a harsh 'churr' note. These can be given in combination, perhaps sounding a bit like song? This is probably NOT what was heard at Hempstead on Sunday but is as close as I can get to a 'female song'. Do west coast birders or museum collection managers know of proven males that show a very weak breast band? Is there evidence for singing by female Varied Thrushes, or other _Zoothera_ thrushes for that matter? Useful Sources: Clement, P. (2000) Thrushes. Princeton University Press. George, T. L. (2002) Varied Thrush. No. 541. Birds of North America. Pyle, P. (1997) Identification Guide to North American Birds. Part 1. Many thanks to Dave Klauber, Andy Guthrie and Julian Hough for specific information and stimulating discussion. I look forward to enlightenment. Unless there is a specific reason, please post to the whole group or let me know if you don't want your thoughts to be shared. Cheers, Angus Wilson New York City http://www.oceanwanderers.com ------------------------------------------------- This mail sent through IMP: http://horde.org/imp/
[ << | >> | ^^ ] Subject: Ageing & Sexing Varied thrushes From: Julian Hough <Jrhough1(AT)AOL.COM> Date: 18 Mar 2003 9:53am Based on my conversations with Angus Wilson re: the NY bird, here's my two pennies worth based on recent CT individuals. (see surfbirds NAmerican Rarities) There is not much precise info available in the popular press, but I think that the best clue to ageing females is the presence of brown tones to the crown, mantle and tail. In good light and close views, these can be seen. The best indicator to look at in the field is a brownish tail for first-winters and a bluish/grey tail for adult females (based on Pyle et al). Also, a moult contrast in the greater coverts may aid ageing, but I think the pattern of replaced feathers may be similar to juv. outers making it difficult to discern differences in pattern between the two generations of feathers. I'd be interested to hear from banders whether this is accurate or not. I think the orange on the underparts may be too variable for ageing (but i don't know for sure). It seems, from my limited field experience and examination of skins that males and females were not too difficult to sex on breast band definition and colour. Basically, first-winter males and adult males are very similar and probably difficult to age in winter. (f-w with brownish tones to the tail and primaries?). Basically, a bird with a blackish breastband, auriculars and dark slate grey mantle should be a male. A bird with pale greyish upperparts, greyish auriculars and breastband is a female. Regarding the NY bird, it looks like a female so maybe females do give a sub-song?? If not, and this is a male then the literature has to be rewritten. Julian Hough, Graphic Designer, Atlas Container Corp.
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Angus Wilson <wilsoa02(AT)MED.NYU.EDU> Date: 18 Mar 2003 9:55am In response to my earlier posting, two regular contributors to ID-F from California sent me their thoughts on the 'singing female' Varied Thrush from Long Island, New York. Both make good points and I hope they don't mind me sharing them with the whole group. Also Dave Klauber, a co-observer of the bird in question, offered his impressions of the vocalizations. I guess one possibility is that the bird is in fact a male (hormonally at least) that for some reason has not developed the appropriate plumage? Another theory, these plausible scenario would be a second (male) Varied Thrush, singing from a grassy knoll just out of sight. Perhaps Oliver Stone can comment on this :) Cheers, Angus Wilson ************************************* Looks like a female to me. I have both sexes wintering in the forest around us this year (a good year for them, locally). The adult males are a nice slate gray in the dark plumage. This bird looks like a Second Year (SY) female from the indistinct breast band and brown dark markings. A SY male would have a more definite breast band. The graphic for this species account in Pyle shows that SY and After Second Year (ASY) birds can be aged and sexed with a high degree of confidence in March. I know nothing about the different races, which appear to be differentiated in the female plumage and breeding range. It appears that we could see all four in a central CA winter. I am only aware of the "vree" song notes and the "tschook" call note. The song often sounds half-hearted (less developed) through winter, and gets more drawn out and buzzier when sung just before they depart from the wintering areas and on the breeding grounds (like in Alaska). I wonder if the females are capable of singing a less- developed song as in the Carpodacus finches? I haven't found any references that say so, however. Les Chibana, Palo Alto CA A couple of very quick thoughts. I have never seen any Varied Thrush lacking a breast band, this is really an odd trait no matter what age or sex you consider the bird to be. However, missing band more likely on younger birds or female, I agree. Darkness of the face and the real blue-grey tone to the back looks good for male. Have no data on singing females in Varied Thrush, I have never seen a female American Robin singing, that is the best I can do. If it was me, I would put the singing behaviour as a clearer sign of sex than the plumage but that is from biases I have. Looking at blackbirds vocal behaviour is very sex dependent with certain vocalizations given by one sex and one sex only, while in others it is variable. Most migratory, sexually dimorphic birds have only singing males. I have banded a few Varied Thrushes and forget how we aged them, other than skull. With thrushes tail feather shape is quite reliable, the youngsters having narrow and pointed tail feathers. It all depends on the moult though, and you probably have looked at Pyle to figure all that out. Alvaro Jaramillo, Alviso, CA I had previously sent my comments to Angus Wilson regarding my thoughts on the Varied Thrush, but I am repeating them per his request to make comments public. I had several brief but decent looks at the Varied Thrush Sunday morning. My thoughts are that it is an immature male, for the following reasons. The dark mask contrasted strongly with the bright and clearly defined orange supercilium, as well as with the orange throat. The clear definition is stronger than any illustrations I checked for female Varied Thrush. I did not see any breast band, which would indicate a female, but the clear and strong definition of the facial pattern makes me think it's a male. Also the bird was singing. I had taped a copy of the song from Peterson's Western bird calls. To my ears there were 2 types: one a long drawn out whistle, and the second similar, but with a more "burry" texture, for lack of a better word. The bird responded at times to the tape, but only with the second "burry" call, which at times was quite loud. The combination of the singing and the strong facial pattern made me believe it was an immature male. Interestingly, I can find no reference as to whether the Varied Thrush is a second or 3rd year bird in the books I have. That includes Sibley, Peterson, National Geo, Golden Guide, the Master Guide series by Audubon, and the old North American Birds with the Fuertes paintings. This last book gives an extremely detailed description, but for adult birds only. Maybe one of the old Bent Life Histories books might have more info. Dave Klauber, NY
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Julian Hough <Jrhough1(AT)AOL.COM> Date: 18 Mar 2003 11:15am Forgive me for sounding flippant,it is no reflection on anybody, but allow me a lighthearted comment.. As someone once said with regards to a two bird theory, "that's one too many!!!" or...maybe its of the confusing subspecies Zoothera Transvestitus!!! Best, Julian Hough, CT, USA
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Tony Leukering <GreatGrayOwl(AT)AOL.COM> Date: 18 Mar 2003 11:17am ----INCLUDING text/plain MIME SECTION---- Hi all: While I have virtually no experience with singing Varied Thrushes (well, actually, none), I will say that there are so many passerine species in which females sing (many more than I would have guessed just a few years ago) that I would be very hesitant to identify the NY VATH as a male, solely on the basis of song (and against the obvious plumage features telling us that the bird is a female). If you think about where most VATHs breed (I'm not talking about that short-statured willow habitat in western Alaska that VATHs call breeding habitat), how many times does one actually see a singing VATH? The species is also relatively little studied, another red flag in making definitive statements such as, "Female VATHs do not sing." George (2000) states a number of times items along this line, "Other aspects of the breeding biology of this species remain poorly known." Also in that reference, he describes aspects of song in the species but nowhere states that females do not sing. In fact, nowhere in the Sounds section is song attributed to males, except that they "respond to taped song playback with countersinging..." and that they "generally sing from top of live conifers; rarely from snags, logs, or ground (Beck 1997)." So, while it may be that males are the only VATHs that sing, I would say that this thesis is far from proven. Literature Cited Beck, M. 1997. reference cited in George (2000). George, T. L. 2000. Varied Thrush (Ixoreus naevius). In The Birds of North America, No. 541 (A. Poole and F. Gill, eds.). The Birds of North America, Inc., Philadelphia, PA. Enjoy, Tony Leukering Brighton, CO ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Les Chibana <les(AT)BIRDNUTZ.COM> Date: 18 Mar 2003 2:00pm I thought that it was hilarious that Alvaro and I sexed this bird differently! In reality, we've seen only a small amount of Varied Thrushes hitting the nets where we've banded. And now that he mentioned it, I don't recall ever seeing a VATH without a breast band either. And on 2nd thought, the facial pattern is a lot darker than I would expect for a SY female that lacks a breast band. I suppose that the bit of gray/grey lesser coverts/scapulars showing between the branches could indicate a SY male or even ASY female. I gave more weight to the lack of breast band, for which I don't have any real basis for except Pyle's account. In reconsidering, I'd have to say that I don't know what age or sex this bird appears to be. I guess I haven't studied them enough. So much for the high degree of confidence rating for ageing and sexing indicated in Pyle. Tony makes a good point. I have only once seen VATH making the call note, and never seen them singing (in CA or AK). So, I can't vouch either way, that males only sing or that females also sing. I suppose that I can't even be so sure that the buzzy "vree" that I hear is a VATH. Les Chibana, Palo Alto CA On Tuesday, March 18, 2003, at 08:54 AM, Angus Wilson wrote: > In response to my earlier posting, two regular contributors to ID-F > from > California sent me their thoughts on the 'singing female' Varied Thrush > from Long Island, New York. Both make good points and I hope they don't > mind me sharing them with the whole group. Also Dave Klauber, a > co-observer of the bird in question, offered his impressions of the > vocalizations. I guess one possibility is that the bird is in fact a > male (hormonally at least) that for some reason has not developed the > appropriate plumage? Another theory, these plausible scenario would be > a > second (male) Varied Thrush, singing from a grassy knoll just out of > sight. Perhaps Oliver Stone can comment on this :) > > Cheers, Angus Wilson > > ************************************* > > Looks like a female to me. I have both sexes wintering in the forest > around us this year (a good year for them, locally). The adult males > are > a nice slate gray in the dark plumage. This bird looks like a Second > Year (SY) female from the indistinct breast band and brown dark > markings. A SY male would have a more definite breast band. The graphic > for this species account in Pyle shows that SY and After Second Year > (ASY) birds can be aged and sexed with a high degree of confidence in > March. I know nothing about the different races, which appear to be > differentiated in the female plumage and breeding range. It appears > that > we could see all four in a central CA winter. > > I am only aware of the "vree" song notes and the "tschook" call note. > The song often sounds half-hearted (less developed) through winter, and > gets more drawn out and buzzier when sung just before they depart from > the wintering areas and on the breeding grounds (like in Alaska). I > wonder if the females are capable of singing a less- developed song as > in the Carpodacus finches? I haven't found any references that say so, > however. > > Les Chibana, Palo Alto CA > > > A couple of very quick thoughts. I have never seen any Varied Thrush > lacking a breast band, this is really an odd trait no matter what age > or > sex you consider the bird to be. However, missing band more likely on > younger birds or female, I agree. Darkness of the face and the real > blue-grey tone to the back looks good for male. Have no data on singing > females in Varied Thrush, I have never seen a female American Robin > singing, that is the best I can do. If it was me, I would put the > singing behaviour as a clearer sign of sex than the plumage but that is > from biases I have. Looking at blackbirds vocal behaviour is very sex > dependent with certain vocalizations given by one sex and one sex only, > while in others it is variable. Most migratory, sexually dimorphic > birds > have only singing males. > > I have banded a few Varied Thrushes and forget how we aged them, other > than skull. With thrushes tail feather shape is quite reliable, the > youngsters having narrow and pointed tail feathers. It all depends on > the moult though, and you probably have looked at Pyle to figure all > that out. > > Alvaro Jaramillo, Alviso, CA > > > I had previously sent my comments to Angus Wilson regarding my thoughts > on the Varied Thrush, but I am repeating them per his request to make > comments public. I had > several brief but decent looks at the Varied Thrush Sunday morning. My > thoughts are that it is an immature male, for the following reasons. > The > dark mask contrasted strongly with > the bright and clearly defined orange supercilium, as well as with the > orange throat. The clear definition is stronger than any illustrations > I > checked for female Varied Thrush. I did not > see any breast band, which would indicate a female, but the clear and > strong definition of the facial pattern makes me think it's a male. > Also > the bird was singing. I had taped a copy > of the song from Peterson's Western bird calls. To my ears there were 2 > types: one a long drawn out whistle, and the second similar, but with a > more "burry" texture, for lack of a > better word. The bird responded at times to the tape, but only with the > second "burry" call, which at times was quite loud. The combination of > the singing and the strong facial > pattern made me believe it was an immature male. Interestingly, I can > find no reference as to whether the Varied Thrush is a second or 3rd > year bird in the books I have. That > includes Sibley, Peterson, National Geo, Golden Guide, the Master Guide > series by Audubon, and the old North American Birds with the Fuertes > paintings. This last book gives > an extremely detailed description, but for adult birds only. Maybe one > of the old Bent Life Histories books might have more info. > > Dave Klauber, NY >
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Mike Patterson <celata(AT)pacifier.com> Date: 18 Mar 2003 3:38pm http://home.pacifier.com/~neawanna/temp/vath.html and females will buzz in the winter... -- Mike Patterson Astoria, OR celata(AT)pacifier.com A child who becomes acquainted with the birds about him hears every sound and puzzles out its meaning with a cleverness that amazes those with ears who hear not. -Neltje Blanchan http://www.pacifier.com/~mpatters/bird/bird.html
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Angus Wilson <wilsoa02(AT)MED.NYU.EDU> Date: 18 Mar 2003 7:46pm Thanks to everyone who has contributed so far. Keep 'em coming! I am forwarding some comments from David Spector who is unable to post to the list. One thing I forgot to mention before about the Long Island Varied Thrush is that someone played a tape once or twice. I don't know whether this might incite a burst of song? Cheers, Angus Wilson ******************************** There are records of female song for many species of temperate zone, migrant species in which normally the males do all of the singing. For example, there are records of rare female song from at least ten North American wood-warblers (I can give you references if you want). Also, females (even in species in which females do not normally sing) can often be induced to sing with testosterone implants; there is no inherent absolute barrier to singing by females in these species. There is the possibility that a bird that is aberrant in one behavior (flying to the East) is more likely to be aberrant in another behavior (singing as a female in a species that normally does not have female song). Thanks for sharing this intriguing bird, David David Spector, New Britain, Connecticut
[ << | >> | ^^ ] Subject: VA Beach Gull From: John Puschock <g_g_allin(AT)HOTMAIL.COM> Date: 18 Mar 2003 9:28pm Martin Reid has posted several of my photos of a gull seen on the March 1 pelagic trip out of Virginia Beach, Virginia. The photos are posted at http://www.martinreid.com/gullsp33.html. The interesting thing about this gull was the relatively light gray of the upperparts and the limited amount of black in the primaries. In direct comparison, it was obviously darker than Herring Gulls -- it wasn't difficult to pick it out of the gull flock following the boat -- but lighter than other Lesser Black-backeds I've seen (all in the eastern U.S.). Assuming the Kodak gray scale printed in the latest issue of Birding is accurate, I'd describe it as a 7 or perhaps slightly lighter. I'd like to hear opinions on the possible identity of this gull. Does it appear to be within the range of Lesser Black-backed Gull or is it better described as a probable Herring x Lesser Black-backed hybrid (or anything else)? For the most part, I'll let the photos speak for themselves -- I spent most of my time trying to get the photos rather than observing the bird. I did lighten the scans a bit to try to match the shade of gray in the original slides, but I didn't make any color corrections. (The film was Fuji Provia 400 and the scans were made on a 13-inch laptop monitor.) On my monitor, the first flight shot looks a little darker than the original slide. In the bottom left photo, the orbital ring appears to be orange-red; however, it appears red in the originals. Leg color was unquestionably yellow -- it didn't strike me as being any different than the leg color of a typical Lesser Black-backed. Also, that's a Herring Gull on the left in the top photo. John Puschock g_g_allin(AT)hotmail.com _________________________________________________________________ The new MSN 8: advanced junk mail protection and 2 months FREE* http://join.msn.com/?page=features/junkmail
[ << | >> | ^^ ] Subject: Identification of Brant - Variability and hybridization From: Angus Wilson <wilsoa02(AT)MED.NYU.EDU> Date: 18 Mar 2003 11:37pm Re: An 'intermediate brant' from New York: photos and discussion Hi everyone, I have put together some pictures of an interesting brant goose taken on Long Island, New York in January of this year. http://www.oceanwanderers.com/IntrmBrantNY.html The bird is one of the puzzling 'intermediate-types' that we've been seeing on a regular basis during the past few years. It was present in a small flock of Low-Arctic Atlantic Brant (aka Pale-bellied Brent Geese) that also included a conventional Black Brant. The latter is now an annual vagrant to the area. At this point, we really don't know what these 'intermediate' birds are exactly but in hopes of stimulating discussion, I review the three leading possibilities. At the bottom of the page there are several links to older web pages showing similar birds from previous winters. Some will be familiar to ID- F regulars but may still be worth revisiting with a fresh eye. As always I look forward to your comments on these interesting and mysterious birds and would urge everyone to share their thoughts or photographs with the entire list. It is clear that there are still many gaps in our knowledge of North American brant. Cheers, Angus Wilson New York City http://www.oceanwanderers.com ------------------------------------------------- This mail sent through IMP: http://horde.org/imp/
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Nathan Hentze <w_tanager(AT)HOTMAIL.COM> Date: 19 Mar 2003 12:31am Hello all, As for the sex of the NY bird I think that it is a male, possibly SY, for reasons stated by others so I won't repeat them again. I am glad that this discussion has arisen, as I have been pondering an individual I observed last year. On Feb. 13, 2002 at my home on eastern Vancouver Island, BC I observed at relatively close range (<15ft) a VATH that in all aspects I could see appeared to be a male (dark slate-blue crown and back, etc) with other VATH of both sexes nearby for comparison. The unusual thing was that save for about 1cm of dark where the breastband should begin, the entire breast was orange. A nearly identical pattern to the bird in question. I have been paying closer attention to my local VATH breastbands since then, and have never seen another individual of any sex that completely lacked it. Females seem to show more variation in completeness of the breastband, with some birds observed having the very centre of the breast with a split of orange feathering, or orange feathering intermixed with the gray-brown band. In another unusual variant I observed this previous weekend, a male had a breastband that was about 2x wider than normal, leaving only the chin and very upper throat orange. So in summary, from my personal observations females show more individual variation in breastband completeness, but males have shown the most extreme variations. I think it is quite plausible that any age or sex of VATH may lack or partially lack a breastband. Thanks for bringing this topic to light, Nathan Hentze, Cumberland, BC, Canada On Tue, 18 Mar 2003 00:32:31 -0500, Angus Wilson <wilsoa02(AT)MED.NYU.EDU> wrote: >Do west coast birders or museum collection managers know of proven males that >show a very weak breast band? Is there evidence for singing by female Varied >Thrushes, or other _Zoothera_ thrushes for that matter? > >Cheers, Angus Wilson >New York City >http://www.oceanwanderers.com
[ << | >> | ^^ ] Subject: Re: VA Beach Gull From: Nick Rossiter <nrossiter(AT)SUPANET.COM> Date: 19 Mar 2003 7:51am Interesting gull with good wingtip shots. It's not likely to be graellsii as there is too little black in the wingtip (P7 has long grey tongues) and too much white (almost complete tip P10, large mirror P9). It's not Azorean or Southern atlantis as there is too little black in the wingtip (P7 has long grey tongues). The amount of white on P9 is also very untypical for these forms. It might be an Iberian atlantis as these can have more strongly indented black on the wingtip and more white on P9. They are slightly darker than michahellis -- around 7 on Kodak grey scale. It's most like cantabricans (Cantabrican Atlantic YLG) from northern Spain/SW France which can be thought of as a Herring Gull (argenteus) in biometrics, calls and wingtip pattern but with slightly darker mantle and finer bill. This form tends slightly paler than michahellis -- hence probably around 5-6 on the Kodak grey scale. Also they can retain head markings well into spring. Ringing shows they are not very mobile. It could of course also be a yellow-legged smithsonianus but the mantle shade seems too dark for this. Another view would be that it's a hybrid graellsii x smithsonianus. Features seem more extreme than intermediate e.g. wing-tip is classical Herring Gull. The leg colour may be critical here. A hybrid would be expected to show some fleshy tones in the legs themselves. As the legs are bright yellow I would discount a hybrid. So I think this is an Atlantic YLG from northern Iberia – typical cantabricans or less typical Iberian atlantis. Cheers … Nick g_g_allin(AT)HOTMAIL.COM wrote: > Martin Reid has posted several of my photos of a gull seen on the March 1 > pelagic trip out of Virginia Beach, Virginia. The photos are posted at > http://www.martinreid.com/gullsp33.html. > > The interesting thing about this gull was the relatively light gray of the > upperparts and the limited amount of black in the primaries. In direct > comparison, it was obviously darker than Herring Gulls -- it wasn't > difficult to pick it out of the gull flock following the boat -- but lighter > than other Lesser Black-backeds I've seen (all in the eastern U.S.). > Assuming the Kodak gray scale printed in the latest issue of Birding is > accurate, I'd describe it as a 7 or perhaps slightly lighter. > > I'd like to hear opinions on the possible identity of this gull. Does it > appear to be within the range of Lesser Black-backed Gull or is it better > described as a probable Herring x Lesser Black-backed hybrid (or anything > else)? > > For the most part, I'll let the photos speak for themselves -- I spent most > of my time trying to get the photos rather than observing the bird. I did > lighten the scans a bit to try to match the shade of gray in the original > slides, but I didn't make any color corrections. (The film was Fuji Provia > 400 and the scans were made on a 13-inch laptop monitor.) On my monitor, > the first flight shot looks a little darker than the original slide. In the > bottom left photo, the orbital ring appears to be orange-red; however, it > appears red in the originals. Leg color was unquestionably yellow -- it > didn't strike me as being any different than the leg color of a typical > Lesser Black-backed. Also, that's a Herring Gull on the left in the top > photo. > > John Puschock > g_g_allin(AT)hotmail.com > > > > > > _________________________________________________________________ > The new MSN 8: advanced junk mail protection and 2 months FREE* > http://join.msn.com/?page=features/junkmail -- Message sent with Supanet E-mail
[ << | >> | ^^ ] Subject: Plumage variation in Varied Thrush From: Julian Hough <Jrhough1(AT)AOL.COM> Date: 19 Mar 2003 10:27am I looked again at Angus Wilson's photos and although I can't see the darkness/grayness of the upperparts, I think that the auriculars, if as dark in life, suggest a male, albeit an atypical one which is totally lacking a breastband! Given the input from observers of the NY bird, and if we assume that males are the only(?) ones to sing, this seems the most logical explanation. I think that Varied Thrush plumages are probably more variable than we think (hence its name??). When you consider that the only Western Palearctic record (Cornwall, England in November 1982) completely lacked any orange pigment in the plumage, a bird lacking a breastband doesn't seem too surprising. Cheers, Julian Hough, CT, USA
[ << | >> | ^^ ] Subject: Re: Plumage variation in Varied Thrush From: Matt Kenne <mkenne(AT)NETAMUMAIL.COM> Date: 19 Mar 2003 11:35am This is a multi-part message in MIME format. ----INCLUDING text/plain MIME SECTION---- You can see a Varied Thrush without a breastband AND without orange = coloration on Don Roberson's thrush family page = http://montereybay.com/creagrus/thrushes.html Matthew Kenne Algona, Iowa mkenne(AT)netamumail.com ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gulls From: Ian Mclaren <iamclar(AT)DAL.CA> Date: 19 Mar 2003 12:45pm All, especially east coasters: I am dealing with a CBC observation at the head of the Bay of Fundy of what were well described by reliable observers (but beginnerish birders) as four Laughing Gulls. These are not unprecedented here in early winter and indeed on a few past CBCs. The description of size, mantle/upperwing tone, and wing tips seem right, althought the birds weren't close enough to show eye-crescents. However, two of the birds had black (blackish?) heads. This isn't supposed to be per literature available. But, does it ever in reality occur? Answer me directly if you think this is not a suitable/interesting topic. Cheers, Ian Ian A. McLaren, PhD Professor Emeritus Phone (W) 902-494-2565 Biology Department Fax 902-494-3736 Dalhousie University Phone (H) 902-429-7024 Halifax, NS B3H 4J1
[ << | >> | ^^ ] Subject: Re: Varied Thrush Puzzle From: Tony Leukering <GreatGrayOwl(AT)AOL.COM> Date: 19 Mar 2003 4:15pm ----INCLUDING text/plain MIME SECTION---- Hi all: I have one more piece of info to provide: In spring 1987 when I was working at American Birds in NYC, I chased a VATH that a friend found at Battery Park (southern tip of Manhattan). The bird was obviously a VATH, but had only a ghost of a breast band. At the time, I assumed that the bird was a female, but it did look dark-crowned and dark-backed - needless to say, I was confused, especially since it was my life VATH. So, though VATHs without (or with greatly reduced) breast bands seem rare, they are not exceedingly so. Perhaps this thread will encourage more careful study of VATHs! Tony Leukering Brighton, CO ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gulls From: Tony Leukering <GreatGrayOwl(AT)AOL.COM> Date: 19 Mar 2003 4:22pm ----INCLUDING text/plain MIME SECTION---- Hi Ian and others: A couple years ago, an adult Franklin's Gull spent the winter here in the Barr Lake, CO, vicinity in complete Alternate plumage - even down to the pink wash on the body. Additionally, I have heard of, but not seen, individual hooded gulls, specifically Bonaparte's, on CBCs. However, I have never seen, nor heard of, a winter Alternate-plumaged Laughing, but I would not at all say that it couldn't happen. However, four together seems exceedingly unlikely. Tony Leukering Brighton, CO ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gulls From: Matt Kenne <mkenne(AT)NETAMUMAIL.COM> Date: 19 Mar 2003 7:28pm This is a multi-part message in MIME format. ----INCLUDING text/plain MIME SECTION---- We've also had a few mid-winter alternate plumage Franklin's Gulls on = Iowa reservoirs, although they were single birds sighted in different = years. I can't say I understand the phenomenon, but it's not a great = leap to project it onto Laughing Gulls. Matthew Kenne Algona, Iowa mkenne(AT)netamumail.com ----- Original Message -----=20 From: Tony Leukering=20 To: BIRDWG01(AT)listserv.arizona.edu=20 Sent: Wednesday, March 19, 2003 5:22 PM Subject: Re: [BIRDWG01] Winter Laughing Gulls Hi Ian and others: A couple years ago, an adult Franklin's Gull spent the winter here in = the Barr Lake, CO, vicinity in complete Alternate plumage - even down to = the pink wash on the body. Additionally, I have heard of, but not seen, = individual hooded gulls, specifically Bonaparte's, on CBCs. However, I = have never seen, nor heard of, a winter Alternate-plumaged Laughing, but = I would not at all say that it couldn't happen. However, four together seems exceedingly unlikely. Tony Leukering Brighton, CO=20 ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: VA Beach Gull From: Martin Reid <upupa(AT)AIRMAIL.NET> Date: 20 Mar 2003 5:05am Dear all, I read with interest Nick Rossiter's comments on the VA gulls. I find a couple of features on this bird puzzling for any form of YLGU: The orbital ring appears yellowish - at best yellowish-orange. Care must be taken when assessing bare parts colors, and John comments that in the original image the orbital ring appears to be red - but the colors of the bill provide a good reference for direct comparison in these two images, and it is hard for me to accept that the gonydal spot appears the normal red color while the orbital ring has somehow been color-shifted - ? LBBG and all forms of YLGU (I think) have red or reddish orbital rings. The pattern of black at the BASE of the primaries; On the upperside there is a small amount of gray visible beyond the primary coverts on both P8 and P9, rather than the black reaching the gray coverts. A close look at a lightened version of the underside of the wing shows P9 and P8 to be almost all-gray from below. The upperside distal pattern of P8; the black on the inner web is very short, and separated from the basal gray by an obvious white "pearl" I know little about Cantabrican Gull, but here are some flight shots: http://www.martinreid.com/mcanp06.html If we remove YLGU from consideration for the sake of argument, what are we left with? I feel that the primary pattern is not typical of the smith HERGs I see in Texas (to little black, and "pearls" too conspicuous") - but in Newfoundland there are smiths that have far less black in the primaries, thus this individual could be a hybrid between such a HERG and a LBBG, I suppose. The strongly yellow legs are odd for an F1 hybrid, and I might expect an F2 (with smith) to look more HERG-like. In some ways, this VA bird reminds me of the dark-backed gull from Texas: http://www.martinreid.com/gullspx4.html - but this TX bird has more flesh tones to the legs, a dark eye, red orbital, etc. My personal feeling on the Texas bird is that it is likely a taimyrensis/birulai hybrid. This form is highly variable in appearance; check out the Ujihara's pages on taim. - most of which are hybrids, I think: http://larus.hp.infoseek.co.jp/a_heuglini.html - these birds have some distinct structure features that seem unlikely in a eastern smith HERG x graellsii/intergrade LBBG. Given that VA is a regular (?) haunt of Black-tailed gull(s), I feel we should consider this possibility! On a related ID gull topic - winter records of small gulls in Alternate plumage - I'd like to add that I see about one or two adult FRGUs a year in mid-winter (late Dec - late Jan) - and they ALL have been in breeding plumage. I've also seen a BOGU on New Year's Day with a complete black hood. If the biological clock is skewed enough for a gull to be at the wrong temporal latitude (not applicable to BOGU, but to LAGU and FRGU) then this may be in response to a mis-timed hormonal trigger, which could also affect their appearance. I think this phenomenon is well-documented in small gulls due to the obvious plumage change - the development of a "black" head - but I wonder if this should also be considered a significant factor in assessing large migratory gulls that appear at the wrong temporal latitude? Just a thought. Good gulling, Martin At 3/19/2003 04:27 AM +0000, you wrote: >Martin Reid has posted several of my photos of a gull seen on the March 1 >pelagic trip out of Virginia Beach, Virginia. The photos are posted at >http://www.martinreid.com/gullsp33.html. > >The interesting thing about this gull was the relatively light gray of the >upperparts and the limited amount of black in the primaries. In direct >comparison, it was obviously darker than Herring Gulls -- it wasn't >difficult to pick it out of the gull flock following the boat -- but lighter >than other Lesser Black-backeds I've seen (all in the eastern U.S.). >Assuming the Kodak gray scale printed in the latest issue of Birding is >accurate, I'd describe it as a 7 or perhaps slightly lighter. > >I'd like to hear opinions on the possible identity of this gull. Does it >appear to be within the range of Lesser Black-backed Gull or is it better >described as a probable Herring x Lesser Black-backed hybrid (or anything >else)? > >For the most part, I'll let the photos speak for themselves -- I spent most >of my time trying to get the photos rather than observing the bird. I did >lighten the scans a bit to try to match the shade of gray in the original >slides, but I didn't make any color corrections. (The film was Fuji Provia >400 and the scans were made on a 13-inch laptop monitor.) On my monitor, >the first flight shot looks a little darker than the original slide. In the >bottom left photo, the orbital ring appears to be orange-red; however, it >appears red in the originals. Leg color was unquestionably yellow -- it >didn't strike me as being any different than the leg color of a typical >Lesser Black-backed. Also, that's a Herring Gull on the left in the top >photo. > >John Puschock >g_g_allin(AT)hotmail.com > > > > > >_________________________________________________________________ >The new MSN 8: advanced junk mail protection and 2 months FREE* >http://join.msn.com/?page=features/junkmail Martin Reid Fort Worth, Texas mailto:upupa(AT)airmail.net http://www.martinreid.com
[ << | >> | ^^ ] Subject: RE : VA Beach Gull From: =?iso-8859-1?Q?Pierre-Andr=E9_CROCHET?= <crochet(AT)CEFE.CNRS-MOP.FR> Date: 20 Mar 2003 6:52am Dear all, I totally agree with Martin that this bird is very puzzling for any form of YLG. The comment's by Nick Rossiter that Cantabrican YLG have wing tip pattern more similar to Herring Gull is not supported by our analyses (Pons et al. in prep) of specimens or our field experience. Cantabrican YLG are typical YLG when in terms of wing tip pattern. There is a slight amount of variation when compared to Mediterranean birds, but it is only in term of frequency of different characters and there is a full overlap. I think the wing tip pattern of this bird is outside the variation of any form of YLG. I might be proved wrong of course, but this is my feeling when studying the pictures. Last, the theory that a large proportion of birds wintering in Japan are hybrids or intergrades between vegae (inc. birulai) and taimyrensis is poorly supported. There seems to be more variation in vegae that most people recognise. The birds breeding in the N. Taimyr peninsula seem to correspond to what some people call taimyrensis~vegae intergrades (pictures can be seen in Filchagov et al. 1992, l'Oiseau et RFO 62:128-148). But the fact that the population breeding there is very homogenous and far from the presumed intergradation zone (SW Taimyr) speaks against this theory. Now, it might be that there has been in the past some intergradation between heuglini/taimyrensis and vegae/birulae and that the characters of the western vegae populations partly derive from this phenomenon. Similarly, the there are strong arguments to suggest that the yellow legged genes of the eastern argentatus populations originate from introgression (from what?). Or that barabensis might have gaine! d some of its morphological characters through past introgression with cachinnans. Still, it does not make them intergrades. The difference is whether there is ongoing gene flow and hybridisation, or whether an ancient event of introgression is involved. Present evidence support the last theory. Finally, I have no idea what the VA Beach Gull can be... Best, Pierre-Andre Pierre-André Crochet CEFE-CNRS 1919, route de Mende 34293 Montpellier cedex 5 France tel: + 33 6 07 32 60 75 fax: + 33 4 67 41 21 38 crochet(AT)cefe.cnrs-mop.fr -----Message d'origine----- De : Martin Reid [mailto:upupa(AT)AIRMAIL.NET] Envoyé : jeudi 20 mars 2003 13:08 À : BIRDWG01(AT)listserv.arizona.edu Objet : Re: [BIRDWG01] VA Beach Gull Dear all, I read with interest Nick Rossiter's comments on the VA gulls. I find a couple of features on this bird puzzling for any form of YLGU: The orbital ring appears yellowish - at best yellowish-orange. Care must be taken when assessing bare parts colors, and John comments that in the original image the orbital ring appears to be red - but the colors of the bill provide a good reference for direct comparison in these two images, and it is hard for me to accept that the gonydal spot appears the normal red color while the orbital ring has somehow been color-shifted - ? LBBG and all forms of YLGU (I think) have red or reddish orbital rings. The pattern of black at the BASE of the primaries; On the upperside there is a small amount of gray visible beyond the primary coverts on both P8 and P9, rather than the black reaching the gray coverts. A close look at a lightened version of the underside of the wing shows P9 and P8 to be almost all-gray from below. The upperside distal pattern of P8; the black on the inner web is very short, and separated from the basal gray by an obvious white "pearl" I know little about Cantabrican Gull, but here are some flight shots: http://www.martinreid.com/mcanp06.html If we remove YLGU from consideration for the sake of argument, what are we left with? I feel that the primary pattern is not typical of the smith HERGs I see in Texas (to little black, and "pearls" too conspicuous") - but in Newfoundland there are smiths that have far less black in the primaries, thus this individual could be a hybrid between such a HERG and a LBBG, I suppose. The strongly yellow legs are odd for an F1 hybrid, and I might expect an F2 (with smith) to look more HERG-like. In some ways, this VA bird reminds me of the dark-backed gull from Texas: http://www.martinreid.com/gullspx4.html - but this TX bird has more flesh tones to the legs, a dark eye, red orbital, etc. My personal feeling on the Texas bird is that it is likely a taimyrensis/birulai hybrid. This form is highly variable in appearance; check out the Ujihara's pages on taim. - most of which are hybrids, I think: http://larus.hp.infoseek.co.jp/a_heuglini.html - these birds have some distinct structure features that seem unlikely in a eastern smith HERG x graellsii/intergrade LBBG. Given that VA is a regular (?) haunt of Black-tailed gull(s), I feel we should consider this possibility! On a related ID gull topic - winter records of small gulls in Alternate plumage - I'd like to add that I see about one or two adult FRGUs a year in mid-winter (late Dec - late Jan) - and they ALL have been in breeding plumage. I've also seen a BOGU on New Year's Day with a complete black hood. If the biological clock is skewed enough for a gull to be at the wrong temporal latitude (not applicable to BOGU, but to LAGU and FRGU) then this may be in response to a mis-timed hormonal trigger, which could also affect their appearance. I think this phenomenon is well-documented in small gulls due to the obvious plumage change - the development of a "black" head - but I wonder if this should also be considered a significant factor in assessing large migratory gulls that appear at the wrong temporal latitude? Just a thought. Good gulling, Martin At 3/19/2003 04:27 AM +0000, you wrote: >Martin Reid has posted several of my photos of a gull seen on the March 1 >pelagic trip out of Virginia Beach, Virginia. The photos are posted at >http://www.martinreid.com/gullsp33.html. > >The interesting thing about this gull was the relatively light gray of the >upperparts and the limited amount of black in the primaries. In direct >comparison, it was obviously darker than Herring Gulls -- it wasn't >difficult to pick it out of the gull flock following the boat -- but lighter >than other Lesser Black-backeds I've seen (all in the eastern U.S.). >Assuming the Kodak gray scale printed in the latest issue of Birding is >accurate, I'd describe it as a 7 or perhaps slightly lighter. > >I'd like to hear opinions on the possible identity of this gull. Does it >appear to be within the range of Lesser Black-backed Gull or is it better >described as a probable Herring x Lesser Black-backed hybrid (or anything >else)? > >For the most part, I'll let the photos speak for themselves -- I spent most >of my time trying to get the photos rather than observing the bird. I did >lighten the scans a bit to try to match the shade of gray in the original >slides, but I didn't make any color corrections. (The film was Fuji Provia >400 and the scans were made on a 13-inch laptop monitor.) On my monitor, >the first flight shot looks a little darker than the original slide. In the >bottom left photo, the orbital ring appears to be orange-red; however, it >appears red in the originals. Leg color was unquestionably yellow -- it >didn't strike me as being any different than the leg color of a typical >Lesser Black-backed. Also, that's a Herring Gull on the left in the top >photo. > >John Puschock >g_g_allin(AT)hotmail.com > > > > > >_________________________________________________________________ >The new MSN 8: advanced junk mail protection and 2 months FREE* >http://join.msn.com/?page=features/junkmail Martin Reid Fort Worth, Texas mailto:upupa(AT)airmail.net http://www.martinreid.com
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gulls From: "D. Heindel" <birdfish(AT)EARTHLINK.NET> Date: 20 Mar 2003 8:24am ----DELETED HTML-ENCODED SECTION----
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gull From: Ian Mclaren <iamclar(AT)DAL.CA> Date: 20 Mar 2003 8:41am Thanks to all who replied personally or on the list. I've decided to list the birds as "Gull sp.," and will perhaps include a comment in my short account of this year's Atlantic Canada CBCs. THere seems to be a consensus that alternate plumage in winter would be very rare or absent in Laughing Gull, but occasional in Franklin's. Although the latter has also been seen here very rarely in early winter, I don't think it can be safely offered as an alternative. Cheers, Ian M. Ian A. McLaren Biology Department Fax 902-494-3736 Dalhousie University Phone (H) 902-429-7024 Halifax, NS B3H 4J1
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gull From: Alvaro Jaramillo <chucao(AT)ATTBI.COM> Date: 20 Mar 2003 9:42am At 11:41 AM 3/20/2003 -0400, Ian Mclaren wrote: >Thanks to all who replied personally or on the list. I've decided to list >the birds as "Gull sp.," and will perhaps include a comment in my short >account of this year's Atlantic Canada CBCs. > >THere seems to be a consensus that alternate plumage in winter would be >very rare or absent in Laughing Gull, but occasional in Franklin's. >Although the latter has also been seen here very rarely in early winter, >I don't think it can be safely offered as an alternative. There are some general patterns in gulls/shorebirds in the wrong plumage for the season. At least I think there are. The key to understanding this is to think of where they winter. Birds that winter in the northern hemisphere are "programmed" to think that spring is coming when day length begins to increase. Birds that winter in the southern hemisphere are "programmed" to think that their spring is coming when day length begins to decrease (because they are thinking spring, when its actually fall in the southern hemisphere!). So northern hemisphere/tropical wintering birds that overshoot and wind up in winter (boreal winter) much farther south may gain a breeding plumage (in boreal winter) if that individual reacts strongly to the day light changes it encounters there. This may explain why a couple of the Short-billed Dowitcher records from Argentina (in boreal winter) were of breeding plumaged birds, and why at least two records of Laughing Gull (in boreal winter) in Chile were of birds in breeding plumage. This doesn't mean that every time a northern hemisphere winterer overshoots it will obtain a messed up plumage, but its more likely to happen in this situation than in any other. Similarly birds that typically winter in the southern hemisphere but for some reason remain in the northern hemisphere may obtain a breeding plumage at the wrong time of the year. If a Franklin's Gull winters in North America, once its internal physiology switches from migration mode to wintering mode it could potentially react to shortening day lengths (in late fall and early winter) and obtain a breeding plumage during the boreal winter. Again, it won't happen every time but its more likely to happen than in other situations. A lot of species (examples - Sanderling, Red Knot, Ruddy Turnstone etc.) winter through a wide latitudinal range, a huge latitudinal range actually. In these cases the pattern I noted above will not apply. But, I bet that if you took a group of Red Knots that winter here in the San Francisco Bay and you released them in Bahia Lomas in southern Patagonia in the fall, some of those birds would switch plumage. Its dependent on what the individual populations are "programmed" to do. I think its a plausible explanation, but entirely untested. Perhaps others can come up with patterns in the birds they know which could show that this seldom applies, or perhaps that it does account for odd plumages they have encountered. So what about the gulls in the Maritimes? Based on this hypothesis, they would be more likely to be Franklin's Gulls than Laughing Gulls - but surely it isn't this simple. cheers, Al Alvaro Jaramillo Biologist San Francisco Bay Bird Observatory P.O. Box 247 Alviso, CA 95002 (408)-946-6548 http://www.sfbbo.org/ chucao(AT)attbi.com
[ << | >> | ^^ ] Subject: Winter LAGU From: Gavin Bieber <kingbird77(AT)HOTMAIL.COM> Date: 20 Mar 2003 10:01am Hello All, A quick note on the Laughing Gull thread. I have seen a few fully hooded Laughing Gulls along the upper Gulf coast (mississippi and alabama) as early as mid Jan. Gavin Bieber Kingbird77(AT)hotmail.com Tucson, AZ _________________________________________________________________ STOP MORE SPAM with the new MSN 8 and get 2 months FREE* http://join.msn.com/?page=features/junkmail
[ << | >> | ^^ ] Subject: Re: VA Beach Gull From: Peter Adriaens <peter.adriaens(AT)DUTCHBIRDING.NL> Date: 20 Mar 2003 10:50am Hi, I tried to post on this bird before, but my message apparently did not get through. Now Martin and Pierre-André have said the most... Perhaps I may contribute in some way by confirming that the primary pattern is indeed quite similar to that of some Newfoundland Herring Gulls (which also means that it is certainly not typical of any Yellow-legged Gull). The extent of its head streaking (with some spots/blotches in the neck) is also unlike Yellow-legged Gull, as is the rather dull bill pattern (the gonys spot is not bright red, and it does not reach on to the upper mandible; there is also a slight blackish band at the gonys). In addition, I would like to point out that several adult hybrid gulls that have bred in Belgium in recent years, have shown quite yellow legs. This includes one colour- ringed bird (ringed by Norman Van Swelm in The Netherlands as a Herring x Yellow-legged Gull). There are no guarantees that leg colour in a hybrid will always be intermediate between that of its parents. Even 'pure' Herring Gulls may show yellow legs. I feel this has to be a hybrid smithsonianus x graellsii. Best regards, Peter Adriaens Belgium
[ << | >> | ^^ ] Subject: VA Beach Gull From: Nick Rossiter <nrossiter(AT)SUPANET.COM> Date: 20 Mar 2003 1:24pm This is a multi-part message in MIME format. ----INCLUDING text/plain MIME SECTION---- I'm not sure that everyone has looked at the cantabricans on Martin = Reid's web site at: http://www.martinreid.com/mcanp06.html This page shows a range of typical cantabricans adults. The wingtips of = these birds are not typical michahellis. True P9 and P10 are similar to = those of Mediterranean michahellis (plenty of white) and hence to = argenteus. But the middle primaries are different. In general these = show less tendency to black on P4, more likelihood of a spot or a thin, = broken band on P5 and a greater tendency to a marked pale grey indent = into the solid black area of the wingtip.=20 My own studies show a similar picture. See=20 http://www.nrossiter.supanet.com/ylg/cantylg.htm for details.=20 In addition the literature abounds with information on the = argenteus-like features of cantabricans including an orange-red orbital = ring and extensive head markings into spring. I will give more details = if required but see Martin Garner & David Quinn, p.58-59 90(1-2), in = their British Birds article from January-February 1997 for a start.=20 I think to some extent we're into classification problems. I only regard = cantabricans as an Atlantic Yellow-legged Gull in the broadest sense. It = does not fit well with the atlantis from further south and I would not = classify it as such. The clasification of cantabricans is very = debatable. It could even be included under argenteus (after all some = argentatus have yellow legs). I think Pierre's DNA studies have linked = it with michahellis.=20 Obviously M. Pons information on cantabricans would be very welcome but = we can only go on the evidence in the public domain. Is there any chance = of making some summary information available perhaps? That would be a = great help.=20 Best wishes ... Nick =20 Citing unpublsihed results does not realy advance the debate unless = you're going to release some information for us to see and evaluate. Is = Mr Pons prepared to do this? I'll call as evidence Martins Reid's site already mentioned in his = message at:...=20 This pattern is much more like a Herring Gull than michahellis. In = Mediterranean michahellis you often get a spot on P4, always a broad = solid band on P5 and a solid black triangle to the wing tip.=20 See Paul Isenmann's seminal paper at http://www.nrossiter.supanet.com/ylg/isenmann.htm and Garner & Quinn (1997).=20 =20 Also see Teyseedre's paper on cantabricans=20 In reply to Martin's comments, cantabricans can show an orange-red = orbital ring. Geographically cantabricans is sandwiched betweem Herring = Gull (argenteus) in western France and Iberian atlantis in north-west = Spain.=20 ----DELETED text/html MIME SECTION----
[ << | >> | ^^ ] Subject: Re: Winter Laughing Gull From: Floyd Hayes <floyd_hayes(AT)YAHOO.COM> Date: 20 Mar 2003 2:10pm An estimated 5,000 Laughing Gulls winter along the west coast of Trinidad. I recall seeing a few winter birds retaining alternate plumage during one winter and just did an electronic search through previous Southeastern Caribbean Bird Alerts. Here's what I found: On 16 December 2000, one of 1500 had a solid black hood (and really surprised me). On 21 January 2001, four of 1567 had a solid black hood (after which many began acquiring alternate plumage). I probably have additional observations in my field notes but don't want to take the time to review them. In any case, the above data illustrate that retention of alternate plumage throughout the winter is quite rare. I don't recall seeing any such birds during the several other winters when I resided in Trinidad. ===== Floyd E. Hayes Wildlife Biologist Division of Fish and Wildlife, 6291 Estate Nazareth, St. Thomas, Virgin Islands 00802, USA Tel: 340-775-6762; Fax: 340-775-3972 Website: http://www.geocities.com/floyd_hayes __________________________________________________ Do you Yahoo!? Yahoo! Platinum - Watch CBS' NCAA March Madness, live on your desktop! http://platinum.yahoo.com
[ << | >> | ^^ ] Subject: Fwd: Re: Winter Laughing Gull From: Alvaro Jaramillo <chucao(AT)ATTBI.COM> Date: 21 Mar 2003 12:07am Folks, Don Roberson sent me this note, and gave me permission to forward it to the group. >Date: Thu, 20 Mar 2003 09:46:29 -0800 >From: creagrus <creagrus(AT)montereybay.com> > >Al -- an excellent discussion on this topic! You wrote: "I think its a >plausible explanation, but entirely untested...." Perhaps it HAS been >'tested' somewhat beyond the annectodol records you mention. The >Monterey Bay Aquarium takes in for their aviary injured shorebirds found >in coastal California. They are held in an aviary that has extensive >light coming through the roof and windows as it faces the ocean, and >presents an 'unnatural' lighting situation. A surprising percentage of >shorebirds there go into full alternate in mid-winter (particularly >Thanksgiving to Xmas): Black-bellied Plover, Dunlin, both species of >dowitchers, Red-necked Phalarope, etc. I've always believed it was the >unnatural lighting and distorted day-length that caused these very odd >molts. > >Cheers, Don Alvaro Jaramillo Biologist San Francisco Bay Bird Observatory P.O. Box 247 Alviso, CA 95002 (408)-946-6548 http://www.sfbbo.org/ chucao(AT)attbi.com
[ << | >> | ^^ ] Subject: RE : VA Beach Gull From: =?iso-8859-1?Q?Pierre-Andr=E9_CROCHET?= <crochet(AT)CEFE.CNRS-MOP.FR> Date: 21 Mar 2003 2:47am This is a multi-part message in MIME format. ----INCLUDING text/plain MIME SECTION---- Hi all, =20 A very short one : yes, there are statistical differences in several = characters of the wing tip pattern between various populations of Yellow = Legged Gulls, including differences between Cantabrican and both = Mediterranean and Southern Atlantic Iberian populations. Yes, there are = some morphological characters of Cantabrican gulls which are similar to = argenteus characters (amount of head streaking in winter, some = morphometric characters). NO, Cantabrican gulls are not similar to = argenteus in wing tip pattern, NO, they are not more similar overall to = argenteus than to michahellis (in fact, when seen side by side, = michahellis and Cantabricans are extremely similar except in winter for = the most streaked individuals of Cantabricans, and an argenteus among = them is really easy to spot: own experience on the Basque Country coast) = and NO they can't be classified as "Herring Gulls". Don't trust the = "literature" too much.=20 =20 The paper with these results by Pons et al. is in revision. We hope to = have it published soon. =20 And last (but this is more personal) NO I don't see how the VA Bach = Gull could be a Cantabrican, or any, YLG.=20 =20 Hope this help, =20 Pierre-Andr=E9 =20 Pierre-Andr=E9 Crochet CEFE-CNRS 1919, route de Mende 34293 Montpellier cedex 5 France tel: + 33 6 07 32 60 75 fax: + 33 4 67 41 21 38 crochet(AT)cefe.cnrs-mop.fr -----Message d'origine----- De : Nick Rossiter [mailto:nrossiter(AT)SUPANET.COM]=20 Envoy=E9 : jeudi 20 mars 2003 21:30 =C0 : BIRDWG01(AT)listserv.arizona.edu Objet : [BIRDWG01] VA Beach Gull =20 I'm not sure that everyone has looked at the cantabricans on Martin = Reid's web site at: http://www.martinreid.com/mcanp06.html = <http://www.martinreid.com/gullsp33.html>=20 =20 This page shows a range of typical cantabricans adults. The wingtips of = these birds are not typical michahellis. True P9 and P10 are similar to = those of Mediterranean michahellis (plenty of white) and hence to = argenteus. But the middle primaries are different. In general these = show less tendency to black on P4, more likelihood of a spot or a thin, = broken band on P5 and a greater tendency to a marked pale grey indent = into the solid black area of the wingtip.=20 =20 My own studies show a similar picture. See=20 http://www.nrossiter.supanet.com/ylg/cantylg.htm for details.=20 =20 In addition the literature abounds with information on the = argenteus-like features of cantabricans including an orange-red orbital = ring and extensive head markings into spring. I will give more details = if required but see Martin Garner & David Quinn, p.58-59 90(1-2), in = their British Birds article from January-February 1997 for a start.=20 =20 I think to some extent we're into classification problems. I only regard = cantabricans as an Atlantic Yellow-legged Gull in the broadest sense. It = does not fit well with the atlantis from further south and I would not = classify it as such. The clasification of cantabricans is very = debatable. It could even be included under argenteus (after all some = argentatus have yellow legs). I think Pierre's DNA studies have linked = it with michahellis.=20 =20 Obviously M. Pons information on cantabricans would be very welcome but = we can only go on the evidence in the public domain. Is there any chance = of making some summary information available perhaps? That would be a = great help.=20 =20 Best wishes ... Nick =20 =20 =20 =20 =20 =20 =20 =20 =20 =20 =20 =20 =20 =20 Citing unpublsihed results does not realy advance the debate unless = you're going to release some information for us to see and evaluate. Is = Mr Pons prepared to do this? =20 I'll call as evidence Martins Reid's site already mentioned in his = message at:...=20 =20 =20 =20 This pattern is much more like a Herring Gull than michahellis. In = Mediterranean michahellis you often get a spot on P4, always a broad = solid band on P5 and a solid black triangle to the wing tip.=20 See Paul Isenmann's seminal paper at http://www.nrossiter.supanet.com/ylg/isenmann.htm and Garner & Quinn (1997).=20 =20 =20 =20 Also see Teyseedre's paper on cantabricans=20 =20 In reply to Martin's comments, cantabricans can show an orange-red = orbital ring. Geographically cantabricans is sandwiched betweem Herring = Gull (argenteus) in western France and Iberian atlantis in north-west = Spain.=20 =20 ----DELETED text/html MIME SECTION----

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